Looks like a grommet to hold a wire or steady the air box. Dont have a Gen 1 so strickly a guess. On the seal question. My 2nd Gen has a seal in the grove, so I would think yours would. If its missing and its a stupid to find part or expensive heres a fix. Small line of RTV sealant on one side. Let it set up some till it skins up some. On the ohter side take petrolium jelly and smear in grove where fit togehter. You may have to use a q-tip or something. Once RTV is skinned up put pieces together. The petrolium jelly should keep it from sticking together next time you want to take apart and make a nice seal. Any extra that squezes out take a razor blade and trim off once set up completly.